This is my Piero with all his main feathers in place rendered in Cycles :
And with the gaps between the wing feathers closed by scaling them in Local X:
Render Time: 3 Minutes 11.37 Seconds in Cycles.
With some final scaling of the tail feathters (for closing gaps) and the wing end feathers (scaled down in width), Piero is now ready for receiving his body feathers in order to complete his feathering: